, , ,

<<


 >>  ()
Pages:     | 1 || 3 |

...

-- [ 2 ] --

.

- (Fragaria viridis, Aconitum barbatum), Cotoneaster melanocarpus, .

, 22 . . , Cotoneaster melanocarpus, : Rosa majalis Spiraea media.

A.l. B.p. ligularietosum glaucae (. 1, . 10-18) , , (, , , 1997;

, 2003).

: Galatella biflora, Artemisia vulgaris, Ligularia glauca, Poa urssulensis, Trisetum sibiricum, Pimpinella saxifraga.

(70-80 ).

, , , .

A.l. B.p. typicum (. 1, . 19-27) Artemisio - Betuletum .

Calamagrostio Betuletum (, 1989). - (, 2003) - (, , , 2010). - .

(, , 2011), (, , , 2011) (, , 2008).

0,4-0,7, Betula pendula, 22 . Populus tremula.

10-60%, Padus avium, Rosa majalis, Salix caprea Frangula alnus. Rubus idaeus, Cotoneaster melanocarpus, Acer negundo.

, 45-100%, 47 77. : Pulmonaria mollis, Brachypodium pinnatum, Rubus saxatilis .

(. 2).

Calamagrostio Betuletum , , , .

Cacalia hastata (. 2, . 1-9) Crepis lyrata, Bistorta major, Erythronium sibiricum, Cacalia hastata, Poa insignis, Melilotoides platycarpos, Equisetum sylvaticum, Senecio nemorensis, Anthriscus sylvestris, Athyrium filix-femina, Veronica longifolia, Veratrum lobelianum.

Calamagrostio arundinaceae Betuletum pendulae Delphinium Cacalia hastata Adonis sibirica retropilosum L07- L08- L10- L10- L08- L09- G-12- L07- L08- L09- L09- L09- G-28- G-05- L09- G-26- G-06- L10- L10- L07- L09- L11- L11- L11- L11- L11- (%):

:

I 3334433 43 333313+23 a1 V VV Betula pendula a1 1 1 + + - I II Populus tremula a1 + - -I Abies sibirica a3 + + + 2 2 I II II Populus tremula I + + +++ ++++++ 2 + ++ ++ b III V III Padus avium I + + +++ +++ + + ++ b I V III Rosa majalis b + + + + + + + + + + + III II I Salix caprea b + + + ++ + + + + I III II Viburnum opulus b ++ + + 1 + + + + - III III Frangula alnus b + + + + + + + - - II Sorbus sibirica Crataegus + + ++ + + b III II II sanguinea b + + + + - I I Rubus idaeus Cotoneaster +++ b I - II melanocarpus Cacalia hastata c +++++++++ +++ + + V II II Crepis lyrata I +++++ ++ + + + + I II V c Bistorta major Erythronium +1++1+++1 + V - I c sibiricum c ++++ ++++ ++V - II Poa insignis I Melilotoides + ++++ +++ + I I c V platycarpos I Equisetum ++++++ + + I c V sylvaticum I + +++++ + I c V Cacalia hastata I Senecio ++++++ - c V nemorensis Anthriscus + + + + + + III - I c sylvestris Athyrium filix + + + + + III - c femina Veronica + + + + II I c longifolia Veratrum + + + + II I c lobelianum Delphinium retropilosum I Lathyrus + + + + + + + + + + + + II II c V pratensis Delphinium + + + + + + + + + + + + II V I c retropilosum I 1 ++++ ++ +1 + I I c V Urtica dioica Picris + ++++++ - V c hieracioides Calamagrostis ++ 1 ++++ I III c epigeios c + +++ ++ - Fragaria viridis Geranium +++++ + - III pratense c c + + +++ + I II I Viola canina c + + +++ I II Ligularia glauca c ++++ - II Achillea asiatica Arctium ++++ - II c tomentosum Aegopodium 3 ++ I I c podagraria Adonis sibirica I + + + + + + + + + + II I c Lathyrus humilis V I 1+ +++++++ - I b V Lonicera tatarica c + ++++++ + I - V Melica nutans c ++++++ ++ - - V Adonis sibirica I + ++++++ I c V Cruciata krylovii ++ +++ - I II b Spiraea media Calamagrostio arundinaceae Betuletum pendulae I Geranium ++ +++++++++++++++++++++ V V c V bifolium I I + ++++++++++++++++++++ + V c V V Viola hirta I ++ ++12+ ++++++++ + + II V V c Poa angustifolia Origanum ++ + + ++ +++ ++++++ + II III V c vulgare Adenophora +++ ++++++ + ++++ ++V II III lilifolia c I Polygonatum + +++ +++ + +I I c V humile c + + + ++ + +I I III Vicia tenuifolia Filipendula ++ +++ ++ - II II c vulgaris Ranunculus ++ + ++ + II I I c polyanthemos Vicio unijugae Pinion sylvestris I +++++++++++ ++++++ + +++++V V c Vicia unijuga V I + + ++++++++++++ ++++ + V III c V Inula salicina I I Lathyrus ++ ++++ + ++ +++++++ ++ III V V pisiformis c I Ptarmica ++++ + ++ + ++ + +++++ + II V c V impatiens Lupinaster +++ ++ +++ +++ +++++ III III III pentaphyllus c Carici macrourae Pinetalia sylvestris c Carex macroura + + + + + + + + 1 1 + 2 2 + 1 1 + + + + + + 1 1V V V I +++1++11++ +++++++ ++1+++V V c V Trollius asiaticus I Heracleum +++++++++ + ++++++ + ++ + V III c V dissectum I + + + + + + + + + + + + 1 2 1 2 + + + III V V c Viola uniflora I Aconitum ++++++ +++ ++ +++ + ++++ V III c V volubile Pteridium 43 +3+ 21++ + + +221 III II III c aquilinum Cimicifuga ++++++ ++ ++++++ III I V c foetida Brachypodio pinnati Betuletea pendulae Pulmonaria ++++++++++++++++++++++++++ V V V c mollis Brachypodium 2+1+22++1++ 111++++1++12+1 V V V c pinnatum I Lilium +++++++++++++++ ++ ++++++ V V V pilosiusculum c c ++++++++++++++++++ + ++++ V V III Vicia sepium c ++++++++1++1+ 1+++ 1 ++1+ V V III Rubus saxatilis Angelica ++++++++++ +++++++ + + ++ V V III sylvestris c I Pleurospermum +++++++++++ +++++ + +++ V III c V uralense I Bupleurum +++++++ ++ +++++ ++++++ V III V aureum c I Calamagrostis 1+++++++1+++1+ 1+ ++ 2 V II c V arundinacea Serratula ++ ++ ++ + +++++++++++ III III V coronata c c ++ + +++++++ + ++++ +++ III III V Iris ruthenica c Agrimonia pilosa + + + + + + + + + + + + + + + + IV V Hieracium +++++ ++ III I umbellatum c Sanguisorba ++++++++++++++++++ +++++++ V V V c officinalis c +++++++++++++++++ ++++++++ V V V Crepis sibirica Ranunculus +1++1+++1+ ++++++++ ++++++ V V V monophyllus c Polygonatum +++++ +++++++++++ ++++ +++ V V V c odoratum I Phlomoides + ++++++++++++++++++++ ++ V V V tuberosa c Thalictrum +++ ++++++++++++ ++++++++ V V V c minus c ++++++++++++++++++ 1 ++++ V V III Galium boreale Dactylis +++++++++++1+ ++++ + ++ + V V III c glomerata c Cirsium setosum + + + + + + + + + + + + + + + + + + + + III V III c Lathyrus gmelinii + 1 + + 1 + + 1 + + + + + + + + + + V I V I +++++++++ ++++++ + ++V II c V Vicia sylvatica I Equisetum ++++++ +++++1++++ V c V pratense c ++ +++++++++++++ I III V Viola mirabilis c + ++++ ++ +2++++++ II II V Lathyrus vernus c +++ ++++++++++ + II V I Elytrigia repens I I Artemisia ++++ +++++ +++++ c V V vulgaris I Polemonium +++ ++++ + +++++ + III I V caeruleum c I I Filipendula ++++ +++ +++++++ c V V ulmaria I Euphorbia + + + + + + + + + + + + + II II V lutescens c Campanula + + + + + + + + + + + II II III c altaica Geranium + + + + + + + + + + + III II II sylvaticum c I + + + + +++++ + - II c V Fragaria vesca Primula ++ + ++++ + ++ I III III cortusoides c c + + + + + + + + + + II I III Galium verum Aconitum + + + + + + + + + II I II c septentrionale Chamaenerion + + + + + + + 1 + II II II c angustifolium Vicia + ++ +++ + + II II II c megalotropis Helictotrichon + + + + ++ ++ I II II c pubescens Campanula + + + + + + + + II II c glomerata Lithospermum + + + ++ ++ - II II c officinale c + + + + + + + III I Veratrum nigrum Taraxacum + + + + + + + II II I officinale c Thalictrum +++ + ++ - II II c simplex Centaurea ++ + ++ - I II scabiosa c Filipendula + ++ ++ - II II stepposa c Solidago + + + + + II - II c virgaurea Veronica + + + + + II I c chamaedrys Alopecurus + + + + + II I I c pratensis Moehringia + + + + + II - I c lateriflora Tanacetum + + + + + II I c vulgare Anemonoides + ++ ++ I I II c caerulea Festuca ++ + + - II I c pratensis c + + + + I I Paris quadrifolia Potentilla + + ++ - I II c chrysantha Potentilla + + + + II I fragarioides c c + + + - I I Galatella biflora Cirsium + + + I - I c serratuloides Hypopitys + ++ I I c monotropa Platanthera + + + I I I c bifolia c + + + - I I Silene repens : Abies sibirica [a1] (23 +), Acer negundo [b] (23 +), Aconitum barbatum [c] (15 +, 25 +), Aconogonon alpinum [c] (11 +), Adoxa moschatellina [c] (7 +, 18 +), Alchemilla vulgaris [c] (23 +, 24 +), Amoria repens [c] (12 +, 13 +), Androsace amurensis [c] (11 +), Artemisia latifolia [c] (10 +), Atragene sibirica [c] (26 +), Betula pubescens [a3] (13 +), Calamagrostis langsdorffii [c] (2 +, 25 +), Caragana frutex [b] (12 3), Carex arnellii [c] (11 +), Carex atherodes [c] ( +), Carex caryophyllea [c] (3 +, 4 +), Carex pallescens [c] (12 +, 13 +), Carex praecox [c] (10 +, 11 +), Carum carvi [c] (12 +), Cenolophium denudatum [c] (11 +), Cerastium davuricum [c] (3 +, 23 +), Cerastium pauciflorum [c] (26 +), Cirsium helenioides [c] (4 +), Conium maculatum [c] (18 +), Corydalis bracteata [c] (7 +), Crepis praemorsa [c] (4 +, 13 +), Cypripedium macranthon [c] (25 +), Dianthus superbus [c] (1 +), Euphorbia discolor [c] (10 +, 25 +), Festuca gigantea [c] (7 +, +), Festuca rubra [c] (13 +), Galeopsis bifida [c] (18 +), Galium mollugo [c] (7 +, +), Geum aleppicum [c] (12 +, 18 +), Geum urbanum [c] (9 +), Glechoma hederacea [c] (11 +, 12 +), Gypsophila altissima [c] (4 +), Hemerocallis minor [c] (2 +, 20 +), Humulus lupulus [c] (17 +, 18 +), Hypericum hirsutum [c] (12 +), Hypericum perforatum [c] (4 +), Impatiens noli-tangere [c] (18 +), Kadenia dubia [c] (12 +), Knautia arvensis [c] (13 +), Lamium album [c] (7 +, 8 +), Leucanthemum vulgare [c] (15 +), Linaria vulgaris [c] (11 +), Lonicera xylosteum [b] (16 +), Lychnis chalcedonica [c] (6 +), Lysimachia vulgaris [c] (13 +), Maianthemum bifolium [c] ( +), Melandrium album [c] (12 +, 13 +), Milium effusum [c] (11 +), Myosotis krylovii [c] (2 +, 7 +), Myosotis nemorosa [c] (4 +), Neottia nidus-avis [c] (25 +), Nepeta pannonica [c] (9 +), Paeonia anomala [c] (2 +, 6 +), Pedicularis sibirica [c] (16 +), Peucedanum morisonii [c] (25 +, 26 +), Phalaroides arundinacea [c] (3 +), Phleum pratense [c] (13 +), Phragmites australis [c] (6 +), Pimpinella saxifraga [c] (12 +, +), Plantago major [c] (12 +), Poa palustris [c] (11 +), Poa trivialis [c] (20 +), Potentilla canescens [c] (12 +), Primula pallasii [c] (13 +), Prunella vulgaris [c] ( +), Pyrola minor [c] (7 +), Ranunculus acris [c] (4 +, 23 +), Ranunculus auricomus [c] (18 +), Ranunculus grandifolius [c] (2 +, 12 +), Ranunculus repens [c] (12 +), Ribes atropurpureum [b] (7 +), Ribes nigrum [b] (9 +, 15 +), Ribes spicatum [b] ( +, 18 +), Rosa acicularis [b] (11 +, 16 +), Sambucus sibirica [b] (11 +), Saussurea controversa [c] (1 +, 25 +), Scrophularia nodosa [c] (3 +), Sedum telephium [c] (4 +, 11 +), Senecio erucifolius [c] (18 +), Seseli libanotis [c] (25 +), Stellaria bungeana [c] (6 +, 11 +), Stellaria graminea [c] (10 +), Thalictrum foetidum [c] (25 +), Trifolium pratense [c] (12 +, 13 +), Trisetum sibiricum [c] (1 +, 6 +), Veronica krylovii [c] (11 +).

. . . 1.

, , .

Delphinium retropilosum (. 2, .

10-19) Lathyrus pratensis, Delphinium retropilosum, Calamagrostis epigeios, Picris hieracioides, Fragaria viridis, Geranium pratens, Urtica dioica, Viola canina, Ligularia glauca, Achillea asiatica, Arctium tomentosum, Aegopodium podagraria.

, (Lathyrus pratensis, Geranium pratense) , . , . .

Adonis sibirica (. 2, . 20-26) Lonicera tatarica Spiraea media. , Lathyrus humilis, Melica nutans, Adonis sibirica, Cruciata krylovii.

- Lathyro - Pinion Brachypodio - Betuletea. : Aconitum septentrionale, Athyrium filix-femina, Lathyrus gmelinii, Milium effusum, Calamagrostis obtusata, Cirsium heterophyllum, Veratrum lobelianum.

: , , , .

Lathyro - Pinion , Trollio asiaticae Populetum tremulae .. (1989) . - (, , , 2010), (, , 2008;

, , , 2011) , Trollio asiaticae Populetum tremulae cruciatetosum krylovii Lashchinsky, Makunina et Gulyaeva 2011, . .

Geranio sylvaticae Pinenion sylvestris. - , : Lathyrus vernus, Aegopodium podagraria, Geranium sylvaticum, Carex arnelli. Lathyrus vernus, Aegopodium podagraria, Geranium sylvaticum , Carex arnelli , (, 2010).

(0,5-0,7) , . 0,3-0, : Padus avium, Viburnum opulus, Salix caprea, Crataegus sanguinea, Ribes spicatum. , 70-100%, 50-90 (. 3).

.

T. a. P. t. cruciatetosum krylovii (. 3, . 1-8) , .

: Crepis lyrata, Cruciata krylovii, Carex cespitosa, Myosotis nemorosa, Ranunculus grandifolius.

, (Carex cespitosa, Myosotis nemorosa), (Crepis lyrata, Cruciata krylovii, Ranunculus grandifolius), , .. (1989).

Trollio asiaticae Populetum tremulae T.a.-P.t. cruciatetosum T.a.-P.t. veratroetosum T.a.-P.t. typicum krylovii nigrum L09- L09- L09- L09- L09- L09- L09- L09- L09- L09- G-36- G-32- G-43- G-37- G-34- G-40- G-45- G-31- L11- L11- L11- L11- L11- L11- L11- L11- L11- (%):

a Betula + 2+ +1+++++111 1 - VV pendula a I I Populus 4 3 4 + 343 43 2433223 4 + 1 VVI tremula a Betula +4 13 + 3 ++ 1 V- pubescens a I Populus + + + + + + 3 V- tremula a I Betula + + + 3 I- pubescens I I + 1 + + ++ +1 +11 +23+++ + 2+ b Padus avium V VV I I Viburnum + ++ ++ + ++ ++ +++ b opulus V V I II + + + ++ +++ + ++ ++ b Rosa majalis I VI I + ++ + + + I b Salix caprea I I Sorbus + + + + + b sibirica III I Crataegus + + + + + III b sanguinea Frangula ++ + + b alnus III Ribes + + ++ b spicatum I Rosa +++ + b acicularis --I T.a.-P.t. cruciatetosum krylovii c Crepis lyrata + + ++ ++++ + + ++VI Carex + + + ++++ c cespitosa V - I Lysimachia + + + + ++ c vulgaris V - I Ranunculus + + +++ c grandifolius V - I Myosotis + + + ++ c nemorosa V- I Cruciata + + ++ + 1 ++++++ c krylovii VI I T.a.-P.t. veratroetosum nigrum c Vicia unijuga + + + + + + + + ++++++++VI V I Veratrum + + + ++++ ++ c nigrum IIV I Peucedanum + + + + + I c morisonii --I I Lupinaster + +++ c pentaphyllus I-I I Filipendula +++ --I c vulgaris Trollio asiaticae Populetum tremulae I I Geranium + ++ ++ ++ + ++++++ +++++ c sylvaticum VVV Lathyrus ++ + ++++++++++++++++ c vernus - VV I Aegopodium 2 1 22 3 3 4 4233554+ c podagraria VV Lathyro gmelinii Pinion sylvestris Aconitum + ++ ++ ++ ++ + ++++++++++++++++ c septentrionale VVV I Lathyrus + + + + + ++ + ++++++++1++++++ VVV c gmelinii I Milium 1+ 1 +++++1+ +++ c effusum - VI I Veratrum + ++ ++ ++ + + +++ c lobelianum VI I II Paris + + + + ++ ++ c quadrifolia III II Cirsium + + + + ++ + c helenioides III I Stellaria + +++ c bungeana I-I Carici macrourae Pinetalia sylvestris Carex + + ++ + ++ ++ + +1++++++++++++3+ c macroura V VV I + + + ++ + ++ +++++++++++++ c Viola uniflora V VV I I Pteridium 2 23 3 +3 3+ 1 122++ 2 33222 V VV c aquilinum I Heracleum + + + + + ++ + +1+++ 1++++++ + c dissectum V VV I Trollius + + ++ ++ ++ ++ + ++ ++++ + ++ VVI c asiaticus I + + ++ ++ ++ + + + + ++++ ++ II Aconitum c VI I volubile II + +++++ +++++ II Cimicifuga c -II foetida Brachypodio pinnati Betuletea pendulae Pulmonaria + + ++ ++ ++ ++ + ++++++++++++++++ c VVV mollis Calamagrosti + + ++ + ++ ++ + ++++++++++++++++ c VVV s arundinacea Agrimonia + + + ++ ++ ++ + ++++++++++++++ c VVV pilosa Brachypodiu 2 1 22 2+ 2+ ++ + +1+ + ++++++++++ c VVV m pinnatum c Vicia sepium + + + + + + + + + + + + + + + + + + + + + + + V V V I Rubus 1 1 ++ + ++ ++ + +++++++ +++ 1+ VVV saxatilis c I + + ++ ++ ++ ++ +++ + +++++++ I Angelica c VI V sylvestris I Bupleurum + + ++ + ++ + ++++ ++++++++++ c VVV aureum I + ++ ++ ++ + ++++++++++++I I Lilium c pilosiusculum VI V I + + ++ ++ ++ + + + +++ + +++ II Serratula c coronata VI I Pleurospermu + + ++ ++ ++ ++ +++ +++ + c m uralense V VI I I + ++ ++ + + ++++ ++ c Iris ruthenica V II I + + + + ++ I Hieracium c umbellatum I II Galium + ++ ++++++++++++++++++++++++ c boreale V VI I I + ++ ++++++++++++ +++++++++++ c Vicia sylvatica V II I + ++ +++++++++++++++ +++++++ I Euphorbia c lutescens II I Equisetum + 1+ +++++++++ +++ +++++ +++ c pratense V VI I I Dactylis + ++ + ++++++++++ +++ ++++++ c glomerata V II I + ++ ++++++ +++++ +++ +++++ I c Crepis sibirica I II Thalictrum + + + ++++++++++++++++++ c minus V VI I I Phlomoides + ++ +++++ + + ++ +++++++++ c tuberosa V II I + ++ +++++++++++ +++ +++ + I Ranunculus c monophyllus I II c Urtica dioica + + ++ 1+1+ 1+111+++ +1+ V VI I I Polygonatum + ++ ++ + + +++ ++++++++ c odoratum V II I ++ + ++++ +++ ++ +++++++ I Polemonium c caeruleum III Filipendula +++ +++++++ ++ + +++++ c ulmaria VVI I I Sanguisorba + + +++ + + + +++++++++ c officinalis VI I I +++ ++ +1+++ ++ +1 + ++ I Equisetum c sylvaticum III Adenophora ++ +++++ + + + ++++ ++ c lilifolia VV I I ++ +++++++ + + + +++++ I I I Cirsium c setosum VI V I Geranium +++ + ++ + ++++ ++++ VI c bifolium V I Anthriscus + ++ ++++++ ++ ++ c sylvestris I VI I +++ + + + ++ + ++ ++ III c Viola hirta VI I I ++ + + + ++++ ++++ VV c Cacalia hastata I + ++ + ++++++++ c Viola mirabilis -IV I I Ptarmica +++ + ++ + + +++++ VI V c impatiens II ++ + + + ++++ ++ III Impatiens noli c tangere III III ++ ++ ++ + + ++ c Elytrigia repens VI I I I ++ +++ ++ + + c Bistorta major V- I I I Chamaenerion ++ + +++ + + ++ c angustifolium VI I I I Delphinium + + +++ + ++ c retropilosum VI I I Paeonia + + ++ ++ ++ c anomala IV I + +++ ++ ++ II Conioselinum c tataricum II I Lathyrus + +++ + + + + VI c pisiformis I I ++ + +++ + II c Melica nutans II I ++ + + ++ + I I Lathyrus c pratensis I II I I Calamagrostis + + ++ + ++ II I c epigeios I +++ +++ + c Inula salicina V- I I Vicia + ++ ++++ V c megalotropis I I Athyrium filix + + + + + ++ c femina II I I Anemonoides + + + +++ c caerulea V- I + + +++ + c Fragaria vesca I II I + + + + + I Glechoma c hederacea - I I +++ + + I Festuca c gigantea - I I +++ + ++ c Galeopsis bifida - II I + + ++ + I Phalaroides c arundinacea II Ranunculus + + + + c repens I I I + ++ + c Poa insignis I I I Carex + ++ ++ c pallescens V - I Maianthemum ++ + + c bifolium I I I Artemisia + + + ++ c vulgaris II I + + 1+ + I Alopecurus c pratensis II Origanum ++ + + c vulgare I-I I + ++ + + I Delphinium c elatum -I I + + + + c Elymus caninus -II Arctium + + + c tomentosum -II I + + + + I Cirsium c heterophyllum I- I + ++ + I Trisetum c sibiricum I- I Adoxa + + + c moschatellina -I Polygonatum + + + c humile I-I I + + ++ I Geranium c pratense I- I + ++ + c Ligularia glauca I-I I Lithospermum + +++ c officinale I-I Taraxacum + + + + c II I officinale I Moehringia + + + + c II lateriflora I Campanula + + + c I- altaica I Senecio ++ + c I- nemorensis I Campanula + + + + c I- I glomerata c Poa angustifolia + + ++ II I c Lamium album + + -I I + + + c Geum rivale -I Hypericum + ++ c hirsutum II Veronica + + + c longifolia II I Thalictrum + + + c simplex I c Viola canina + + + I-I Tanacetum + + + c vulgare II Ranunculus + + c acris III Cirsium + + + c serratuloides II Veronica + + + c chamaedrys II : Achillea asiatica [c] (8 +, 22 +), Amoria repens [c] ( +), Bromopsis inermis [c] (5 +), Calamagrostis langsdorffii [c] (15 +), Cardamine impatiens [c] (14 +), Carex atherodes [c] (4 +), Cerastium davuricum [c] (10 +), Circaea alpina [c] (12 +), Crepis praemorsa [c] (2 +), Cypripedium calceolus [c] ( +), Dryopteris carthusiana [c] (11 +, 15 +), Dryopteris dilatata [c] (26 +), Equisetum arvense [c] (12 +), Erythronium sibiricum [c] (4 +), Euphorbia discolor [c] (12 +, +), Festuca pratensis [c] (2 +), Galium verum [c] (4 +), Geum aleppicum [c] (26 +), Helictotrichon pubescens [c] (2 +), Kadenia dubia [c] (3 +, 7 +), Lamium album [c] (9 +, 12 +), Lonicera xylosteum [b] (16 +), Lychnis chalcedonica [c] (4 +, 7 +), Matteuccia struthiopteris [c] (3 +, 11 +), Melandrium album [c] (2 +, 10 +), Melilotoides platycarpos [c] (4 +), Myosotis arvensis [c] (2 +, 6 +), Myosotis krylovii [c] (13 +), Neottia nidus-avis [c] (3 +), Oberna behen [c] (2 +, 22 +), Phragmites australis [c] (4 +, 15 +), Picris hieracioides [c] (1 +, 2 +), Platanthera bifolia [c] ( +, 3 +), Poa palustris [c] (3 +, 10 +), Poa trivialis [c] (11 +), Potentilla fragarioides [c] (4 +), Primula cortusoides [c] (27 +), Prunella vulgaris [c] (2 +), Ranunculus acris [c] (14 +, 26 +), Ranunculus auricomus [c] (3 +), Ranunculus polyanthemos [c] (2 +), Rubus idaeus [b] (10 +), Solanum kitagawae [b] (6 +), Spiraea media [b] (22 +, 24 +), Trommsdorffia maculata [c] (3 +), Vicia tenuifolia [c] (10 +, 27 +).

. . . 1.

T. a. P. t. veratroetosum nigrum (. 3, . 19-27) L11-37, , , (55,01449 .. 86,13966 ..), 03.06.2011. : .. .

: Vicia unijuga, Veratrum nigrum, Peucedanum morisonii, Lupinaster pentaphyllus, Filipendula vulgaris.

Lathyro Pinion Vicio Pinion (, 2012). , (0,5-0,7) 20 . . Padus avium. , 55-100%.

T. a. P. t. typicum (. 3, . 9-18) .

Lathyro Pinion, , Campanulo trachelium Betuletum pendulae (, , 2011).

: Erythronium sibiricum, Anemonoides altaica, Corydalis bracteata, Adenophora lilifolia, Bistorta major, Campanula trachelium, Anemonoides caerulea.

Campanulo trachelium Betuletum pendulae L08- L09- L10- L09- L09- L08- L08- L08- L08- L08- G-64- G-62- G-63- G-66- G-67- G-59- G-70- L08- G-60- G-56- (%):

a1 3454 31+ IV Populus tremula a1 24+34+3+23+1111 1 + 3 1 2 V V Betula pendula a1 + I Larix sibirica a1 1 I Pinus sylvestris a3 + I Abies sibirica b ++11+++++++ + + + + IV IV Padus avium b +++ ++++ + + + + II IV Sorbus sibirica b ++ ++++ + + + III II Salix caprea b +++ + + + + + - IV Malus baccata b + ++ + + + + II II Viburnum opulus Campanulo trachelium Betuletum pendulae c Adenophora lilifolia + + + + + + + + + + + + + + + + + III V Campanula trachelium c ++++++++++++ + + + + + IV V c + ++ +++++ IV Bistorta major Erythronium c 1+1+++1++1 + V I sibiricum Anemonoides c 1+1 +++11 IV altaica c Corydalis bracteata 2 + + + 1 + III Anemonoides c + ++ + + + III caerulea Lathyrus pratensis c + + ++ + + + + + + + I V Poa angustifolia c ++ + + + + + + + - V Achillea asiatica c Lathyrus pratensis + + + + + + + + + - V Taraxacum c + + + + + + + + - IV officinale c +++ + + + + + - IV Festuca pratensis c + ++ + + + + - IV Galium mollugo Lathyro gmelinii Pinion sylvestris c + + ++ 1+ + ++ ++ ++ + + + + + + + V V Lathyrus gmelinii Aconitum c + ++ ++ + + ++ ++ + + + + + + IV V septentrionale c + + + ++ + 21 ++ ++ + + + + + V IV Milium effusum Veratrum c + + ++ + + ++ + + + + + IV III lobelianum Cirsium helenioides c + + + + ++ + + + + + III III Athyrium filix c ++ + + + + + + III II femina c + + ++ + + + IV Paris quadrifolia Carici macrourae Pinetalia sylvestris c + ++ ++ + ++ ++ ++ + + + + + + + V V Aconitum volubile Heracleum c + + ++ ++ + + +1 ++ 1 1 1 1 1 + V V dissectum Pteridium aquilinum c 1 +2 + 1 3+ 41 21 2 1 + + + + 2 IV V c 2 1 ++ + ++ ++ ++ + + + + + + + IV V Carex macroura c 1 + 1+ ++ + ++ + + + + V II Trollius asiaticus c + + ++ 1+ + IV Viola uniflora Brachypodio pinnati Betuletea pendulae c Bupleurum aureum + + + + + + + + + 1 + + + + + + + + + + V V c 11 1+ ++ + + ++ ++ + + + + + + V V Rubus saxatilis c Lilium pilosiusculum + + + + + + + + + + + + + + + + + + V V Pleurospermum c ++ ++ + + ++ ++ ++ + + + + + + V V uralense c ++ ++ ++ + ++ ++ + + + + + + V IV Pulmonaria mollis c ++ ++ ++ + ++ + ++ + + + + + V IV Vicia sepium Brachypodium c 1 3 + 1+ 2 ++ 1+ + + + + + V III pinnatum Calamagrostis c 1 + + + +1 ++ + + + + + + IV IV arundinacea c + + + + + + + + + + I IV Agrimonia pilosa c + + + + + + ++ + V Angelica sylvestris c + + + + II Serratula coronata c + + + II Iris ruthenica Polemonium c + + ++ ++ + ++ ++ ++ + + + + + + + V V caeruleum Ranunculus c + ++ ++ + +1 1+ ++ + + + + + + + V V monophyllus c + ++ + + ++ ++ ++ + + + + + + + IV V Cacalia hastata c + + ++ ++ + ++ + ++ + + + + + V IV Vicia sylvatica Euphorbia c + + + + + + + + + + + + + + + + + V IV lutescens c + + + + 1 + + + + + + + + + + + V III Galium boreale c + + + + + + + + + + + + + + + + IV IV Crepis lyrata c + + + + + + + + 1 + + + 1 1 + 1 IV V Festuca gigantea c + + + + + + + + + + + + + + + + V IV Urtica dioica c + + + + + + + + + + + + + + + + V IV Poa insignis c 1 + + + + + + + + + + + + + + + V IV Crepis sibirica c + + + + + + + + + + + + + + + + IV IV Filipendula ulmaria Anthriscus sylvestris c 1 + + ++ ++ + + + + + + + + IV IV Geranium c + + + + + + ++ + + + + + + III IV sylvaticum Polygonatum odoratum c ++++ ++ ++ + + + + + + III V Veronica c + ++ + + + ++ + + + + + + III IV chamaedrys c + + + ++ + ++ + + + + + III IV Dactylis glomerata Impatiens noli c + ++ + ++ + + + + + + + III IV tangere Equisetum c + + + ++ ++ + + + + + + II V sylvaticum c + + + + ++ + + + + + + II V Fragaria vesca Aegopodium c + 2 + 3 1 1 1 2 + 2 1 2 II IV podagraria c + + + + + + + + + + + I V Ptarmica impatiens c + + + + + + + + + + + II IV Vicia unijuga c + + + + + + + + + + V Thalictrum minus Ranunculus c + ++ + + + + + + + - V polyanthemos Delphinium c + ++ + + + + + + - V retropilosum c ++ + ++ + ++ + V Lamium album Adoxa moschatellina c + ++ ++ + ++ + V II c ++ + ++ + + + I I Cirsium setosum I V c + + ++ + + + + I Cruciata krylovii I c + +++ + + + + I V Geum rivale I V c + + + + + + + + Carex pallescens I Arctium c + + + + + + + + - V tomentosum II I c + + + + + + + + I Delphinium elatum Sanguisorba c ++ + ++ + + IV officinalis II Equisetum I pratense c + + + + + + + I II c + + + + + + - I Tanacetum vulgare c ++ + + + + III Artemisia vulgaris c + + + ++ + III Myosotis krylovii II c + + + + + + III I Myosotis nemorosa Phlomoides tuberosa c ++ + ++ + III II Cardamine c + + + + + + - I impatiens II Ranunculus I grandifolius c + + + + + + I II Moehringia c + + + + + + - I lateriflora II I c + + + + + Angelica decurrens c + + + + + III Geranium bifolium Chamaenerion c + + + + II angustifolium Calamagrostis c ++ + + II epigeios c + + + + - II Poa palustris Senecio nemorensis c + + + + II c 2 1 + + II Lathyrus vernus c + + + + II Saussurea latifolia c + + + + II Stachys sylvatica Maianthemum c + + + II bifolium Hypericum hirsutum c ++ + II c + + + II Paeonia anomala c + + + II Viola hirta Scrophularia c + + + II nodosa c + + + I I Veronica longifolia Conioselinum c + + + II tataricum c + + + II Galeopsis bifida : Abies sibirica [a3] (5 +), Betula pubescens [a3] (2 +), Calamagrostis langsdorffii [c] (10 +), Calamagrostis obtusata [c] (3 +), Campanula altaica [c] (1 +, 2 +), Carex muricata [c] (2 +), Cimicifuga foetida [c] (10 +), Cirsium serratuloides [c] (4 +, 6 +), Dactylorhiza fuchsii [c] (18 +, 20 +), Dryopteris filix-mas [c] (2 +, 3 +), Elymus caninus [c] (2 +), Elytrigia repens [c] (2 +), Epilobium montanum [c] (5 +), Geranium albiflorum [c] (8 +), Geranium krylovii [c] (4 +), Larix sibirica [a1] (10 +), Linaria vulgaris [c] (2 +), Listera ovata [c] (3 +), Lysimachia vulgaris [c] (3 +, 5 +), Matteuccia struthiopteris [c] (3 +, 18 +), Melica nutans [c] (6 +, 10 +), Melilotoides platycarpos [c] (8 +), Oberna behen [c] (2 +, +), Origanum vulgare [c] (10 +), Pimpinella saxifraga [c] (2 +), Pinus sylvestris [a1] (10 1), Poa trivialis [c] (18 +), Populus tremula [a3] (14 +), Ranunculus repens [c] ( +), Ribes atropurpureum [b] (3 +), Rosa majalis [b] (2 +, 5 +), Rubus idaeus [b] ( +), Stellaria bungeana [c] (5 +), Stellaria graminea [c] (17 +), Veratrum nigrum [c] (7 +, 11 +), Vicia megalotropis [c] (2 +).

. . . 1.

18 , , , , . , .

, , . Pteridium aquilinum Anthriscus sylvestris Brachypodium pinnatum.

, (, , 2011). , (. 4).

Lathyrus pratensis, : Poa angustifolia, Achillea asiatica, Lathyrus pratensis, Taraxacum officinale, Galium mollugo, Festuca pratensis.

, Molinio-Arrhenatheretea R. Tx. 1937 em. R. Tx.

1970, .

, . , , .

- , , , . Carex macroura Calamagrostis arundinacea. Brachypodio - Betuletea (Bupleurum aureum, Angelica sylvestris) Carici macrourae - Pinetalia (Viola uniflora, Pteridium aquilinum, Trollius asiaticus). sylvestris Calamagrostio epigei-Betuletalia pendulae Heracleum sibiricum, . Peucedano morisonii-Betulion pendulae.

, , Calamagrostio -Betuletalia Peucedano-Betulion, (, , , 1991).

- Primulo cortusoidis - Betuletum pendulae (. 5).

: Polygonatum humile, Spiraea media, Viola mirabilis, Carex obtusata, Veronica longifolia, Primula cortusoides, Ligularia glauca, Aconitum barbatum.

. , - - . , Carici - Pinetalia, (, , , 1997). , Carex obtusata Veronica longifolia, .

Primulo cortusoidis - Betuletum pendulae 1 2 3 4 5 6 7 8 9 10* 11 12 13 14 15 16 17 18 19 20 G-13- G-17- G-16- G-18- G-19- G-14- G-15- G-12- G-11- L11- L11- L11- L11- L11- L11- L11- L11- L11- L11- L11- L11- , (%):

70 70 70 60 60 70 70 70 60 40 60 60 50 60 70 70 60 60 60 60 20 20 40 30 30 20 40 60 10 70 50 70 70 60 70 50 10 20 50 40 60 50 40 70 60 50 60 40 60 40 30 45 40 50 30 50 55 60 40 60 61 42 47 46 46 42 49 62 50 67 72 56 53 54 57 62 71 63 52 54 a1 111+2144+242424442 2 4 V Betula pendula a1 2+1 ++++3313+4+111 + 2 V Populus tremula a3 +2+2+++++2 2 III Populus tremula b +++++++1+41+++++1++ + 1 V Padus avium b +++++++++1++++++++ + + V Rosa majalis b +++++++++ + + III Crataegus sanguinea b 332322 2 3 II Caragana arborescens b 4++++++ + II Ribes nigrum b +++++ + II Viburnum opulus b +++ + I Malus baccata b ++ + I Lonicera tatarica b ++ + I Ribes spicatum b ++ + I Solanum kitagawae Primulo cortusoidis - Betuletum pendulae c ++++++2+21++++++++1 + + V Polygonatum humile b ++++++++122343+1+ + + V Spiraea media c ++++++++++++++++++ + V Viola mirabilis c +++++++1+11++1+1+ + V Carex obtusata c +++++++++++ ++++ + + V Veronica longifolia c ++++++++++ +++++ + IV Primula cortusoides c +++ +++ ++++ + III Ligularia glauca c +++++++ + II Aconitum barbatum Peucedanum morisoni-Betulion pendulae c ++++++++++++++++++ + + V Fragaria viridis c ++++++++++++++++++ + + V Filipendula stepposa c +++++++++++++++++ + + V Phlomoides tuberosa c +++++++++ ++++ + IV Galium verum c ++ ++++++++++++ IV Peucedanum morisonii c + +++++++++ + III Filipendula vulgaris C +++++ +++ II Helictotrichon pubescens c +++++ + + II Carex praecox c +++++++ II Artemisia latifolia c ++ ++ + II Seseli libanotis c Ranunculus polyanthemos + + + + + II c +++ I Polygonatum odoratum Carici macrourae-Pinetalia sylvestris c ++++++++ + + + III Aconitum volubile Calamagrostio epigei-Betuletalia pendulae c ++++++++1++++++++1 + + + V Poa angustifolia c ++++++++++++++++++ + + + V Geranium bifolium c ++++++++++++++++ + + + V Calamagrostis epigeios c +++++++++++++++++ + + V Kadenia dubia c ++++++++++ III Artemisia macrantha c ++ +++ ++++ III Galatella biflora Brachypodio pinnati-Betuletea pendulae c +++++++1+111++1111 + + 1 V Rubus saxatilis c ++++++++++++++++++ + + + V Lilium pilosiusculum c +++++++++++++++++ + + + V Iris ruthenica c ++++++++++++++++++ + + V Pleurospermum uralense c +++++ +++++++++++ + + + V Serratula coronata c ++++++++++++++++ + + + V Agrimonia pilosa c ++ +++1++++1++++ + + + V Brachypodium pinnatum c +++++++++++++++ + + V Pulmonaria mollis c ++ + ++ II Hieracium umbellatum Molinio-Arrhenatheretea c +++++++++++++++++ + + + V Sanguisorba officinalis c +++++++++++++++++ + + + V Thalictrum simplex c +++++++++++++++++ + + + V Elytrigia repens c ++ +++++++++ + + + IV Taraxacum officinale c ++ +++++++++ + + IV Tanacetum vulgare c ++++++++ +++ + + IV Achillea asiatica c +++++++++++ III Lathyrus pratensis c + + +++ II Alopecurus pratensis c +++ + + II Plantago media c ++ + I Geranium pratense c ++++++++++++++++++ + + + V Galium boreale c ++++++++++++++++++ + + + V Vicia cracca c ++++++++++++++++++ + + V Adenophora lilifolia c ++++++++++++++++ + + 1 V Maianthemum bifolium c ++++++++++++++++ + + V Sedum telephium c +++++++++++++ + + + + V Viola canina c + ++++++++++++ + + + IV Lupinaster pentaphyllus c ++++++++++++++ + + IV Lathyrus pisiformis C + + + + + + + + + + + + + IV Cirsium setosum c + + + + + + + + + + + + III Silene repens c + + + + + + + + + + + + III Urtica dioica c + + + + + + + + + + + + III Moehringia lateriflora c + + + + + + + II Solidago virgaurea c + + + + + + II Chamaenerion angustifolium c + + + + + + II Veratrum nigrum c + + + + + + II Lysimachia vulgaris c + + + + + + II Crepis sibirica c + + + + + + II Vicia unijuga c + + + + + + II Carex caryophyllea c + + + + + II Viola hirta c + + + + + II Campanula altaica c + + + + + II Lathyrus vernus c + + + + + II Carex pediformis c + + + + I Poa palustris c + + + + I Lithospermum officinale c + + + 2 I Equisetum pratense c + + + I Athyrium filix-femina c + + + I Paris quadrifolia c + + + I Adonis vernalis c + + + I Geranium sylvaticum c + + + I Sonchus arvensis c + + + I Ranunculus auricomus c + + + I Melandrium album c + + + I Calamagrostis canescens : Acer negundo [b] (11 +, 14 +), Allium nutans [c] (6 +), Anemone sylvestris [c] (13 +), Artemisia dracunculus [c] (17 +), Betula pendula [a3] (12 +), Botrychium multifidum [c] (21 +), Bromopsis inermis [c] (9 +, 20 +), Cacalia hastata [c] (6 +), Campanula glomerata [c] (8 +, 9 +), Caragana frutex [b] (19 +), Carex cespitosa [c] (11 +), Carex macroura [c] (8 +), Centaurea scabiosa [c] (8 +), Cirsium serratuloides [c] (15 +, 17 +), Cirsium vulgare [c] (15 +), Crepis praemorsa [c] (16 +, 20 +), Cypripedium calceolus [c] (16 +), Cypripedium guttatum [c] (16 +), Dactylis glomerata [c] (20 +), Dianthus superbus [c] (11 +, 17 +), Dryopteris dilatata [c] (16 +), Euphorbia virgata [c] (3 +, 4 +), Festuca pratensis [c] (20 +), Festuca pseudovina [c] (13 +), Fragaria vesca [c] (7 +), Frangula alnus [b] (15 +), Geum aleppicum [c] (8 +, 16 +), Glechoma hederacea [c] (3 +, 19 +), Heracleum dissectum [c] (8 +, 16 +), Hierochloe sibirica [c] (14 +, 16 +), Hypopitys monotropa [c] (20 +), Inula salicina [c] (15 +, 16 +), Lathyrus humilis [c] (9 +), Leonurus tataricus [c] (9 +, 20 +), Linaria vulgaris [c] (9 +), Matteuccia struthiopteris [c] (14 +, 16 +), Orthilia secunda [c] (16 +), Oxytropis campanulata [c] (13 +), Plantago urvillei [c] (13 +), Poa insignis [c] (8 +), Ranunculus monophyllus [c] (16 +, 17 +), Rumex thyrsiflorus [c] (11 +, 17 +), Salix caprea [b] (14 +), Sambucus sibirica [b] (19 +), Scutellaria galericulata [c] (9 +), Senecio erucifolius [c] (16 +), Silene nutans [c] (15 +), Solanum kitagawae [c] (19 +), Thalictrum foetidum [c] (13 +), Thalictrum minus [c] (7 +, 19 +), Thalictrum petaloideum [c] (7 +, 19 +), Veronica krylovii [c] (13 +, +), Vicia megalotropis [c] (9 +), Vicia sepium [c] (7 +, 19 +), Viola arenaria [c] ( +).

. . . 1.

. , . 16-20 0.5 0.7. . (40-60% ), , Caragana arborescens. , , 40-60%, 40-60 . Poa angustifolia Rubus saxatilis Carex obtusata Polygonatum humile. . .

Calamagrostio - Betuletalia. , , , .

, , , , .. (1950) .

, (, , 2007). , - , . (, , 2012).

.

Betula pubescens, Carex elongata Carex cespitosa. 30%.

: Carex elongata, Poa palustris, Carex riparia, Festuca rubra.

, 0,7-0,9 14-16 . , 40 50 , 60-70%. Carex cespitosa, Carex riparia, Phragmites australis Filipendula ulmaria (. 6).

Carici elongatae-Betuletum albae Alnetea glutinosae, .. (2009) .

- , Milio-Abietea. , (, 1932). , Milio-Abietea Saussureo latifoliae-Populetum tremulae, .. (2003).

Saussureo latifoliae Populetum tremulae L10- L10- L09- G-20- L10- L10- L10- L10- L10- L10- (%):

40 70 70 50 70 70 60 60 60 - - 10 20 50 50 50 30 30 90 100 100 85 75 80 90 85 80 : 34 40 34 40 36 47 43 48 44 a1 1 2 + 2 1 III Abies sibirica a1 3 4 + 2 4 4 4 3 4 3 V Populus tremula a3 + + + + + III Abies sibirica b + + + + + + + + + + V Ribes atropurpureum b + + + + + 2 + 1 + V Padus avium b + + 3 2 + 2 1 1 IV Caragana arborescens b + + + + II Sambucus sibirica b + + + II Sorbus sibirica b + + + II Viburnum opulus Galium odoratum c + + + + + + 1 + + V Galium odoratum c + + + + + + + + IV Conioselinum tataricum Saussureo latifoliae Populetum tremulae c 4 5 4 + 1 3 3 3 1 V Matteuccia struthiopteris c + + + + + + + IV Allium microdictyon c + + + + + + + IV Saussurea latifolia Milio-Abietea c + + + + + + + + + + V Stellaria bungeana c + + + + + + + 1 + + V Urtica dioica c + + + + 1 + + + 1 1 V Aconitum septentrionale c 1 + + + + + + + + V Anemonoides altaica c + + + + + + + + + V Paris quadrifolia c + + + + + + + + + V Euphorbia lutescens c + + + + + + + + + V Milium effusum c + + + + + + + + IV Corydalis bracteata c + + + + + + + + IV Lathyrus gmelinii c 1 + + + 1 + + IV Erythronium sibiricum c + + + + + + + IV Myosotis krylovii c + + + + + + + IV Viola uniflora c + + + + II Geranium krylovii c + + + II Dryopteris carthusiana c + + + + II Paeonia anomala c + + + + II Cirsium helenioides c + + + + + + + + + + V Lamium album . c + + + + + + + + + + V Anemonoides caerulea c + + 1 + + + + + + V Heracleum dissectum c + + + + + + + + + V Ranunculus monophyllus c + + + + + + + + + V Crepis sibirica c + + + + + + + + + V Athyrium filix-femina c + + + + + + + IV Humulus lupulus c + + + + + + + IV Adoxa moschatellina c + + + + + + III Polemonium caeruleum c + + + + + + III Pteridium aquilinum c + + + + + + III Delphinium elatum c + + + + + + III Anthriscus sylvestris c + + + + + + III Filipendula ulmaria c + + + + + III Calamagrostis langsdorffii c + + + + + III Pulmonaria mollis c + + + + + III Impatiens noli-tangere c + + + + + III Pleurospermum uralense c + + + + + III Bupleurum aureum c + + + + + III Stachys sylvatica c 1 + 1 + 1 III Aegopodium podagraria c + + + + II Veratrum lobelianum c + + + + II Geum urbanum c + + + + II Thalictrum minus c + + + II Gagea granulosa : Actaea spicata [c] (2 +), Alfredia cernua [c] (5 +, 9 +), Angelica decurrens [c] (4 +, 8 +), Angelica sylvestris [c] (4 +, 7 +), Asarum europaeum [c] (3 +), Bromopsis benekenii [c] (3 +), Calamagrostis obtusata [c] ( +), Carduus crispus [c] (5 +), Daphne mezereum [b] (6 +), Diplazium sibiricum [c] ( +, 7 +), Dryopteris dilatata [c] (3 +), Dryopteris filix-mas [c] (3 +), Festuca gigantea [c] (9 +, 10 +), Glechoma hederacea [c] (5 +), Hesperis sibirica [c] (6 +), Lilium pilosiusculum [c] (2 +, 8 +), Lonicera xylosteum [b] (6 +, 8 +), Oxalis acetosella [c] (3 +, 7 +), Phalaroides arundinacea [c] (8 +), Picea obovata [a3] (6 +), Pinus sibirica [a3] (6 +), Spiraea media [b] (8 +), Tilia sibirica [a3] (3 +), Vicia sepium [c] (2 +, 8 +), Vicia sylvatica [c] (2 +, 8 +).


. . . 1.

, . , Padus avium Sorbus sibirica. Matteuccia struthiopteris ( 120 ). . , , , Galium odoratum.

: Matteuccia struthiopteris, Allium microdictyon, Saussurea latifolia, Conioselinum tataricum, Galium odoratum.

Galium odoratum, Galium odoratum .

Saussureo -Populetum (, , , , 2011).

(. 7). , , , , , .

1 2 3 4 5 6 7 8 9 10 11 12 Betula pendula V IV V V V IV III V V III II I Populus tremula V II II II II IV II IV II III V Betula pubescens I I II I II V I II II Pinus sylvestris I II I Abies sibirica I I II Populus tremula III II I I I II I I I III II I I Betula pubescens I I II I Spiraea media V II I II II I I I I I I Salix caprea I I I II I II III II I III II II I Padus avium V I II III III IV III V IV III IV IV V Viburnum opulus II I II II III II IV II IV III II III Rosa majalis V III III III IV III I III II II I I Crataegus sanguinea III II I I II II I I I I Sorbus sibirica I I III II II II IV II II Ribes spicatum I I I I II I II II I Frangula alnus I I I II III I I II I Lonicera tatarica I I I I III II I I Rubus idaeus I I I I I I II Rosa acicularis I I I I I II I Ribes nigrum II I I I I Sambucus sibirica I I I I II Cotoneaster melanocarpus III I I II II Malus baccata I II II Caragana arborescens I I II I I II Acer negundo I I Ribes atropurpureum I IV II Solanum kitagawae I V II II Polygonatum humile I IV I I II I I Viola mirabilis V IV IV IV V III I II IV I I Carex obtusata V Veronica longifolia V I I I II I II I I I Primula cortusoides IV III IV III II III II I I Ligularia glauca III II III I III I I II I Aconitum barbatum II IV II II I I I I Peucedanum morisonii III III IV IV II I II I Filipendula vulgaris III IV III III II II I II I Pedicularis sibirica IV I I Trommsdorffia maculata III I I I Tephroseris integrifolia III I Thalictrum foetidum I III I I I Plantago urvillei I III I Pulsatilla patens III I I Myosotis imitata III I I Veronica spicata II I Bupleurum multinerve II Oxytropis campanulata I II Artemisia vulgaris I III I I III IV II I I II I Pimpinella saxifraga II II I I I I I Trisetum sibiricum I I I III I Poa urssulensis II I I Silene nutans I III I I Veratrum nigrum II V IV V I III I III I I I I Adonis sibirica II II II IV I I Cruciata krylovii II I I IV I I I III IV III II I Melica nutans II I I IV I I II II I II I Lathyrus humilis I V II I IV I II I Delphinium retropilosum I II II I IV II I II III III I Lathyrus pratensis III II II I I IV II I I III IV I Urtica dioica III II II I IV II IV III III IV IV V Picris hieracioides I I III I II I Geranium pratense I II II I III III Viola canina IV II II II I III I I II I I Arctium tomentosum I I I III I I I IV I I Achillea asiatica III IV II II II I I I I III Erythronium sibiricum I I I V I I I IV IV Crepis lyrata I I I I III V II II V V IV II Poa insignis I II I I II I V II I II III III II Bistorta major II I II II II IV II III I II Equisetum sylvaticum I I I IV IV II IV IV III II Melilotoides platycarpos I II I I I III I I I I Cacalia hastata I I I I III III I III V III V Senecio nemorensis I I III I II I III II Anthriscus sylvestris I I I I III IV III I IV IV IV Origanum vulgare IV III III IV III II I II II I Viola hirta I IV III IV IV V IV II II III I Vicia tenuifolia II II II II I I I I Adenophora lilifolia V IV IV V IV II V I III IV III III IV I Carex cespitosa I I III I I IV I I Lysimachia vulgaris I I I I I III II I Myosotis nemorosa I I I II I I III III I Ranunculus grandifolius I Geranium sylvaticum I I II II II II IV IV III III V III I II V II III IV III Aegopodium podagraria IV II II III IV V II II IV V Lathyrus vernus I II I I I I I IV Galium mollugo II I III Festuca pratensis I I I I I II III Taraxacum officinale III III II II I II II I I I IV II Campanula trachelium I I I I II I IV I III Anemonoides caerulea IV I III Anemonoides altaica III I I I II Corydalis bracteata III I I I II Matteuccia struthiopteris I IV I II Saussurea latifolia III I Allium microdictyon II I I I II III II Conioselinum tataricum III Galium odoratum III Lathyro gmelinii-Pinion sylvestris I II II II II II V V IV IV V Aconitum septentrionale V III III III V II V V V III V V Lathyrus gmelinii V I I I II II II IV II III Veratrum lobelianum II I I II II II I I IV Paris quadrifolia I IV I I I II II I III III Cirsium helenioides III I II II II I III II Athyrium filix-femina I IV I I I V II V IV Milium effusum V I I I II I II Stellaria bungeana V I I III Cirsium heterophyllum I Calamagrostis obtusata II Vicio unijugae-Pinion sylvestris Inula salicina I IV IV IV III IV III I I IV I I Vicia unijuga II V V IV IV V V I V IV IV II I IV Lathyrus pisiformis IV V IV IV IV IV IV I I IV V IV III III III III II Lupinaster pentaphyllus I I I III III IV V III IV I II IV V I Ptarmica impatiens III I II I I I Saussurea controversa Calamagrostio epigei-Betuletalia pendulae V II IV II II Calamagrostis epigeios I II II II I II I Geranium bifolium V IV IV V V V IV I IV IV I II Poa angustifolia V IV III II IV IV II II II I V I II II Kadenia dubia VI I I Galatella biflora II IV IV II I I Artemisia macrantha II II II I Carici macrourae-Pinetalia sylvestris Aconitum volubile III III III III III III V II III V V IV II Heracleum dissectum I I IV III III IV V IV IV III V IV IV Carex macroura I V V V V V V V V IV V IV II II III III III III III IV IV IV V Pteridium aquilinum IV III IV III IV IV V V V II V III Trollius asiaticus V II IV III V V IV IV IV V II I Viola uniflora III III IV II II V I III III III Cimicifuga foetida I I Brachypodio pinnati-Betuletea pendulae Brachypodium pinnatum V V V V V V V IV V V IV V I Lilium pilosiusculum V IV V V V IV V III V III V IV II Pulmonaria mollis V IV V V V V V V V V IV V IV Vicia sepium I IV III IV IV V V IV V IV IV IV II Agrimonia pilosa V IV IV IV V V I IV IV IV IV II I Pleurospermum uralense V II IV IV III IV V IV II V IV IV II Serratula coronata V V IV III IV IV III II III V I II Rubus saxatilis V V V V IV V V V III IV V IV II III III IV III IV IV V III V V Bupleurum aureum III I III IV III V V III IV V I IV Angelica sylvestris II V V V IV V IV III II III III Iris ruthenica I I Hieracium umbellatum II V II II II III I II II I V V V III IV V V V IV IV IV Calamagrostis arundinacea Galium boreale V V V V IV V V V V V IV IV I Crepis sibirica II V IV IV V V V IV IV V IV V V Ranunculus monophyllus I III III III V V V IV III V V V V Dactylis glomerata I IV IV IV III IV V IV IV IV II III II Thalictrum minus I V V V V IV V IV IV II I IV II Filipendula ulmaria I II II I IV IV III IV IV IV IV III Sanguisorba officinalis V V V V V V V II V III I II Vicia sylvatica II II II III III V V V V IV IV II Polygonatum odoratum I V IV IV IV V V II IV III III III I Euphorbia lutescens II III III II IV II V IV V IV IV V Campanula altaica I IV I II III II II I I II I I Vicia megalotropis I III II II II II II I IV I I I Fragaria vesca I II II II IV III I II II I IV II Elytrigia repens V II III II I V II II II III I Polemonium caeruleum II II I III III IV III III III V IV Maianthemum bifolium V I I I I I I II I I I Equisetum pratense I II II V IV III IV V IV I I Chamaenerion angustifolium II II II I II II I II III II I Paeonia anomala I I I I I III I I I II Cirsium setosum III I IV III III V III II III IV III I I Solidago virgaurea II III II II I II I I I I Fragaria viridis V III III II III I I Phlomoides tuberosa V V V IV IV V V III V V I III I Cirsium serratuloides I II II I I I I I I I Alopecurus pratensis II I I I I II I III I I Lithospermum officinale I III II I II II I II I Tanacetum vulgare III I I I II I II III I Filipendula stepposa V IV III II II II I Impatiens noli-tangere I I I II II II III II II Campanula glomerata I III II II II II I II I Veronica chamaedrys I I II II II II III II I Poa palustris I I I I I I II I I Sedum telephium IV I I I I I I I Hypericum hirsutum I I I II I II II I Potentilla fragarioides II I I I II II I I Lamium album I I I I I I III V Calamagrostis langsdorffii I I I I I II II Thalictrum simplex V I II I II II II Euphorbia discolor II I I I I I I Adoxa moschatellina I I I I II IV IV Glechoma hederacea I I I I I II I Galium verum III V IV II II II I I I Helictotrichon pubescens II IV II II II II I I I Carex praecox II I I I II Artemisia latifolia II IV II II I Ranunculus polyanthemos II V II II I II II I I III Seseli libanotis II III II II I I Anemone sylvestris I II I I Galeopsis bifida I I I II I I Centaurea scabiosa I IV II I II I Phalaroides arundinacea I I I III I I Silene repens III II I I I II Trifolium pratense II I I II I I Hemerocallis minor I II I I I I Ranunculus acris I I I I I II Linaria vulgaris I II I I II I Myosotis krylovii I I I I II III Angelica decurrens I I II II I Geum rivale I II III I I Ranunculus repens I I II I I Potentilla chrysantha II I I II I Festuca gigantea I II IV IV III Veronica krylovii I II II I I Aconogonon alpinum I I II I Humulus lupulus I I I III Myosotis arvensis I I I II Vicia cracca V I I I Delphinium elatum II III I IV Carex pallescens I II III IV Platanthera bifolia I I I II Lychnis chalcedonica I I II I Brachypodium sylvaticum I I II Geranium krylovii I I II Carex caryophyllea II I I Dryopteris carthusiana I I II Gypsophila altissima I I I Astragalus danicus I I II Stachys sylvatica I II III Festuca valesiaca I I Alfredia cernua I II Carex pediformis II I Oxalis acetosella I II Medicago falcata I I Dracocephalum ruyschiana I I II Dryopteris filix-mas I II Cardamine impatiens I II Plantago media II Artemisia austriaca II Polygala comosa II Phleum phleoides II . 1-13 - : 1 - . Primulo cortusoidis-Betuletum pendulae;

2 - c. Artemisio latifoliae Betuletum pendulae talictrietosum foetidum;

3 - . Artemisio latifoliae Betuletum pendulae ligularietosum glaucae;

4 - . Artemisio latifoliae Betuletum pendulae typicum;

5 - .

Calamagrostio arundinaceae - Betuletum pendulae Adonis sibirica;

6 .

Calamagrostio arundinaceae - Betuletum pendulae Delphinium retropilosum;

7 - . Calamagrostio arundinaceae - Betuletum pendulae Cacalia hastata;

8 - . Trollio asiaticae-Populetum tremulae typicum;

9 c. Trollio asiaticae-Populetum tremulae veratrosum nigrum;

10 - .

Trollio asiaticae-Populetum tremulae cruceatetosum krylovii;

11 - . Campanulo trachelium Betuletum pendulae Lathyrus pratensis;

12 - . Campanulo trachelium Betuletum pendulae;

13 - . Saussureo latifoliae Populetum tremulae.

, . , II . , , , .

5. .

.

, , .

, . , , (, , , 2011)

( ., 2011).

- (, 2011). .

, . . .

5.1.

. (Livingston, 1903), . (Clements, 1905). . , , , ( , 1983). , .

1928 , , , (Clements, 1928). (, 1997;

, 1999;

, 2002, 2004;



Pages:     | 1 || 3 |
 
 >>  ()





 
<<     |    
2013 www.libed.ru - -

, .
, , , , 1-2 .